Peer-reviewed papers using SHARP data

Sayers, CJ, MR Roeder, L Forrette; D Roche, GLB Dupont, SE Apgar, AR Kocek*, AM Cook, WG Shriver, CS Elphick, B Olsen, and DN Bonter. Accepted. Geographic variation of mercury in breeding tidal marsh sparrows of the northeastern United States. Ecotoxicology.

Roeder, MR, CE Hill, CS Elphick, M Conway, AR Kocek, A Tegeler, and S Woltmann. Accepted. Genetic data disagree with described subspecies ranges for Seaside Sparrow on the Atlantic coast. Condor: Ornithological Applications.

Klingbeil, BT, JB Cohen, MD Correll, CR Field, TP Hodgman, AI Kovach, EE Lentz, BJ Olsen, WG Shriver, WA Wiest, and CS Elphick. 2021. High uncertainty over the future of tidal marsh birds under current sea-level rise projections. Biodiversity and Conservation 30:431-443. [pdf]

Ladin, ZS, WA Wiest, MD Correll, EL Tymkiw, M Conway, BJ Olsen, CS Elphick, WL Thompson, and WG Shriver. 2020. Detection of local-scale  population declines through optimized tidal marsh bird monitoring design. Global Ecology and Conservation 23:e01128. [pdf]

Marshall, H, EJ Blomberg, V Watson, M Conway, JB Cohen, MD Correll, CS Elphick, TP Hodgman, AR Kocek, AI Kovach, WG Shriver, WA Wiest, and BJ Olsen. 2020. Habitat openness and edge avoidance predict Saltmarsh Sparrow abundance better than habitat area. Condor. [pdf]

Field, CR, and CS Elphick. 2019. Quantifying the return on investment of social and ecological data for conservation planning. Environmental Research Letters 14:124081. [pdf]

Walsh, J, GV Clucas, MD MacManes, EK Thomas, and AI Kovach. 2019. Divergent selection and drift shape the genomes of two avian sister species spanning a saline–freshwater ecotone. Ecology and Evolution 9:13477-13494. [pdf]

Field, CR, KJ Ruskin, JB Cohen, TP Hodgman, AI Kovach, BJ Olsen, WG Shriver, and CS Elphick. 2019. Framework for quantifying population responses to disturbance reveals that coastal birds are highly resilient to hurricanes. Ecology Letters 22:2039-2048. [pdf] Featured in the New York Times here.

Conway, M, and BJ Olsen. 2019. Contrasting drivers of diversification rates on islands and continents across three passerine families. Proceedings of the Royal Society, B 286:20191757. [pdf]

Billerman, SM, and J Walsh. 2019. Historical DNA as a tool to address key questions in avian biology and evolution: A review of methods, challenges, applications, and future directions. Molecular Ecology Resources 19:1115-1130. [pdf]

Steen, VA, CS Elphick, and MW Tingley. 2019. An evaluation of stringent filtering to improve species distribution models from citizen science data. Diversity and Distributions 25:1857-1869. [pdf]

Olsen, BJ, JL Froehly, AC Borowske, CS Elphick, CR Field, AR Kocek, AI Kovach, RA Longnecker, WG Shriver, J Walsh, and KJ Ruskin. 2019. A test of a corollary of Allen’s rule suggests a role for population density. Journal of Avian Biology 50:e02116. [pdf]

Walsh, J, PM Benham, PE Deane-Coe, P Arcese, BG Butcher, YL Chan, ZA Cheviron, CS Elphick, AI Kovach, BJ Olsen, WG Shriver, VL Winder, and IJ Lovette. 2019. Genomics of rapid ecological divergence and parallel adaptation in four tidal marsh sparrows. Evolution Letters 3:324-338. [pdf]

Roberts, SG, RA Kern, MA Etterson, CS Elphick, BJ Olsen, and WG Shriver. 2019. Preventing local extinctions to tidal marsh endemic sparrows in Eastern North America. Condor: Ornithological Applications 121:duy024. [pdf] Featured by The Wildlife Society here and in the Philadelphia Inquirer here.

Correll, MD, CS Elphick, W Hantson, TP Hodgman, BB Cline, EL Tymkiw, WG Shriver, and BJ Olsen. 2019. A multi-scale comparison of elevation measurement methods in northeastern tidal marshes of the United States. Wetlands 39:633-643. [pdf]

Correll, MD, W Hantson, TP Hodgman, BB Cline, CS Elphick, WG Shriver, EL Tymkiw, and BJ Olsen. 2019. Fine-scale mapping of coastal plant communities in the northeastern USA. Wetlands 39:17-28. [pdf]

Wiest, WA, MD Correll, BG Marcot, BJ Olsen, CS Elphick, TP Hodgman, GR Gunterspergen, and WG Shriver. 2019. Estimates of tidal marsh bird densities using Bayesian networks. Journal of Wildlife Management 3:109-120. [pdf]

Klingbeil, BT, JB Cohen, MD Correll, CR Field, TP Hodgman, AI Kovach, BJ Olsen, WG Shriver, WA Wiest, and CS Elphick. 2018. A focal species for tidal marsh bird conservation in the northeastern United States. Condor: Ornithological Applications 120:874-884. [pdf] [blog post]

Longenecker, RA, JL Bowman, B Olsen, SG Roberts, CS Elphick, P Castelli, and WG Shriver. 2018. Short-term resilience of New Jersey tidal marshes to Hurricane Sandy. Wetlands 38:565-575. [pdf]

Greenlaw, JS, CS Elphick, W Post, and JD Rising. 2018. Saltmarsh Sparrow (Ammodramus caudacutus), version 2.0. In The Birds of North America (PG Rodewald, ed). Cornell Lab of Ornithology.

Borowske, A, CR Field, KJ Ruskin, and CS Elphick. 2018. Consequences of breeding system for body condition and survival throughout the annual cycle of tidal marsh sparrows. Journal of Avian Biology 49:jav-01529. [pdf]

Benvenuti, B, J Walsh, KM O’Brien, and AI Kovach. 2018. Plasticity in nesting adaptations of a tidal marsh endemic. Ecology & Evolution doi:10.1002/ece3.4528 [pdf]

Walsh, J, AI Kovach, BJ Olsen, WG Shriver, and I Lovette. 2018. Bidirectional adaptive introgression between two ecologically divergent sparrow species. Evolution [pdf]

Walsh, J, LM Maxwell, and AI Kovach. 2018. The role of mating strategy, reproductive success, and mate compatibility in maintaining species boundaries in an avian hybrid zone. Auk: Ornithological Advances 135:693-705. [pdf]

Benvenuti, B, J Walsh, KM O’Brien, M Ducey and AI Kovach. 2018. Annual variation in offspring sex ratios of Saltmarsh Sparrow supports Fisher’s hypothesis. Auk: Ornithological Advances 135:342-358. [pdf]

Field, CR, KJ Ruskin, B Benvenuti, A Borowske, JB Cohen, L Garey, TP Hodgman, RA Kern, E King, AR Kocek, AI Kovach, KM O’Brien, BJ Olsen, N Pau, SG Roberts, E Shelly, WG Shriver, J Walsh, and CS Elphick. 2018. Quantifying the importance of geographic replication and representativeness when estimating demographic rates, using a coastal species as a case study. Ecography 41:971-981. [pdf]

Correll, MD, WA Wiest, TP Hodgman, JT Kelley, BJ McGill, CS Elphick, WG Shriver, ME Conway, CF Field, and BJ Olsen. 2018. A Pleistocene disturbance event explains modern diversity patterns in tidal marsh birds. Ecography 41:684-694. [pdf]

Field, CR, AA Dayer, and CS Elphick. 2017. Landowner behavior can determine the success of conservation strategies for ecosystem migration under sea-level rise. Proceedings of the National Academy of Sciences 114:9134-9139. [pdf]  [blog post]

Ruskin, KJ, MA Etterson, TP Hodgman, AC Borowske, JB Cohen, CS Elphick, CR Field, RA Longenecker, E King, AR Kocek, AI Kovach, KM O’Brien, N Pau, WG Shriver, J Walsh, and BJ Olsen. 2017. Demographic analysis demonstrates systematic but independent spatial variation in abiotic and biotic stressors across 59 percent of a global species range. Auk: Ornithological Advances 134:903-916. [pdf] [blog post]

Roberts, SG, RA Kern, MA Etterson, KJ Ruskin, CS Elphick, BJ Olsen, and WG Shriver. 2017. Factors that influence Seaside and Saltmarsh sparrow vital rates in coastal New Jersey, USA. Journal of Field Ornithology 88:115-131. [pdf]

Walsh, J, IJ Lovette, V Winder, CS Elphick, BJ Olsen, WG Shriver, and AI Kovach. 2017. Subspecies delineation amid phenotypic, geographic, and genetic discordance in a songbird. Molecular Ecology 26:1242-1255. [pdf]

Walsh, J, WG Shriver, MD Correll, BJ Olsen, CS Elphick, TP Hodgman, RJ Rowe, KM O’Brien, and AI Kovach. 2017. Temporal shifts in the saltmarsh-Nelsons’s sparrow hybrid zone revealed by population surveys and genetic data. Conservation Genetics 18:453-466. [pdf]

Nightingale, J, and CS Elphick. 2017. Tidal flooding is associated with lower ectoparasite intensity in nests of the Saltmarsh Sparrow Ammodramus caudacutus. Wilson Journal of Ornithology 129:122-130. [pdf]

Borowske, AC, C Gjerdrum, and CS Elphick. 2017. Timing of migration and prebasic molt in tidal marsh sparrows with different breeding strategies: comparisons among sexes and species. Auk: Ornithological Advances 134:51-64. [pdf]

Field, CR, T Bayard, C Gjerdrum, JM Hill, S Meiman, and CS Elphick. 2017. High-resolution tide projections reveal extinction threshold in response to sea-level rise. Global Change Biology 35:2058-2070. [pdf]

Correll, MD, WA Wiest, TP Hodgman, WG Shriver, CS Elphick, BJ McGill, K O’Brien, and BJ Olsen. 2017. Predictors of specialist avifaunal decline in coastal marshes. Conservation Biology 31:172-182. [pdf] Featured in Audubon Magazine here.

Ruskin, KJ, MA Etterson, TP Hodgman, A Borowske, JB Cohen, CS Elphick, CR Field, RA Kern, E King, AR Kocek, AI Kovach, KM O’Brien, N Pau, WG Shriver, J Walsh, and BJ Olsen. 2017. Seasonal fecundity is not related to geographic position across a species’ global range despite a central peak in abundance. Oecologia 183:291-301. [pdf]

Field, CR, C Gjerdrum, and CS Elphick. 2016. How does choice of statistical method to adjust counts for imperfect detection affect inferences about animal abundance? Methods in Ecology and Evolution 7:1282-1290. [pdf] [data]

Correll, MD, WA Wiest, BJ Olsen, WG Shriver, CS Elphick, TP Hodgman. 2016. Habitat specialization explains avian persistence in tidal marshes. Ecosphere 7(11):e01506. 10.1002/ecs2.1506. [pdf]

Field, CR, C Gjerdrum, and CS Elphick. 2016. Forest resistance to sea-level rise prevents landward migration of tidal marsh. Biological Conservation 201:363-369. [pdf]

Walsh, J., B.J. Olsen, K.J. Ruskin, W.G. Shriver, K.M. O’Brien, and A.I. Kovach. 2016. Extrinsic and intrinsic factors influence fitness in an avian hybrid zone. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12837 [pdf]

Walsh, J., W.G. Shriver, B.J. Olsen, and A.I. Kovach. 2016. Differential introgression and the maintenance of species boundaries in an advanced generation avian hybrid zone. BMC Evolutionary Biology 16:65, DOI 10.1186/s12862-016-0635-y [pdf]

Wiest, WA, MD Correll, BJ Olsen, CS Elphick, TP Hodgman, DR Curson, and WG. Shriver. 2016. Population estimates for tidal marsh birds of high conservation concern in the northeastern USA from a design-based survey. Condor: Ornithological Applications 118:274-288. [pdf]

Ruskin, KJ, MA Etterson, TP Hodgman, and BJ Olsen. 2015. Divergent oviposition preferences of sister species are not driven by nest survival: the evidence for neutrality. Behavioral Ecology and Sociobiology, 69(10):1639-1647. Doi: 10.1007/s00265-015-1975-0 [pdf]

Walsh, J., R.J. Rowe, B.J. Olsen, W.G. Shriver, and A.I. Kovach. 2015 Genotype-environment associations support a mosaic hybrid zone between two tidal marsh birds. Ecology & Evolution, doi: 10.1002/ece3.1864 [pdf]

Walsh, J, WG Shriver, BJ Olsen, KM O’Brien, AI Kovach. 2015. Relationship of phenotypic variation and genetic admixture in the Saltmarsh–Nelson’s sparrow hybrid zone. Auk: Ornithological Advances 132(3):704-716. DOI: 10.1642/AUK-14-299.1 [pdf]

Kovach, AI, J Walsh, J Ramsdell, and K Thomas. 2015. Development of diagnostic microsatellite markers from whole genome sequences of Ammodramus sparrows for assessing admixture in a hybrid zone. Ecology and Evolution, doi: 10.1002/ece3.1514 [pdf]

Elphick, CS, S Meiman, and MA Rubega. 2015. Tidal-flow restoration provides little nesting habitat for a globally vulnerable saltmarsh bird. Restoration Ecology 23:439–446. [pdf] [cover article]

Elphick, CS, and CR Field. 2014. Monitoring indicators of climate change along long Island Sound: A simple protocol for collecting baseline data on marsh migration. Wetland Science and Practice 31:7-9. [pdf]

Wiest, WA, WG Shriver, and KD Messer. 2014. Incorporating climate change with conservation planning: a case study for tidal marsh bird conservation in Delaware, USA. Journal of Conservation Planning 10:25–42. [pdf]


  • Bayard, T. 2010. Testing the role of social cues in saltmarsh sparrow habitat selection decisions. PhD Dissertation. University of Connecticut, Storrs, CT. [pdf]
  • Benvenuti, B. 2016. Nesting adaptations and conservation of a threatened tidal-marsh-nesting bird, the Saltmarsh Sparrow. MS Thesis. University of New Hampshire, Durham, NH. [pdf]
  • Borowske, AC. 2015.  Effects of life history strategies on annual events
    and processes in the lives of tidal marsh sparrows. PhD Dissertation.  University of Connecticut, Storrs, CT. [pdf]
  • Correll, M. 2015. The biogeography and conservation of tidal marsh bird communities across a changing landscape. PhD Dissertation. University of Maine, Orono, ME. [pdf]
  • Field, CR. 2016. A threatened ecosystem in a human-dominated landscape: tidal marsh conservation in the face of sea-level rise. PhD Dissertation.  University of Connecticut, Storrs, CT. [pdf]
  • Garey, LK. 2017. Community-structure and food-web dynamics in northesatern U.S. tidal marshes. MS Thesis. University of Maine, Orono, ME. [pdf]
  • Hill, JM. 2008. Postfledging movement behavior and habitat use of adult female saltmarsh sharp-tailed sparrows. MS Thesis. University of Connecticut, Storrs, CT. [pdf]
  • Kern, RA. 2010. Secretive marsh bird response to prescribed fire in mid-Atlantic tidal marshes. MS Thesis. University of Delaware, Newark, DE. [pdf]
  • Kern, RA. 2015. Conservation ecology of tidal marsh sparrows in New Jersey. PhD Dissertation. University of Delaware, Newark, DE. [pdf]
  • Maxwell, LM. 2018. Drivers of introgression and fitness in the saltmarsh-Nelson’s sparrow hybrid zone. MS Thesis. University of New Hampshire, Durham, NH. [pdf]
  • Meiman, ST. 2011. Modeling saltmarsh sparrow distribution in Connecticut. MS Thesis. University of Connecticut, Storrs, CT. [pdf]
  • Roberts, SG. 2016. Population viability of seaside and saltmarsh sparrows in New Jersey. MS Thesis. University of Delaware, Newark, DE. [pdf]
  • Ruskin, KJ. 2015. Intra- and interspecific variation in demographic rates and niche across the range of a species, the Saltmarsh Sparrow (Ammodramus caudcutus). PhD Dissertation. University of Maine, Orono, ME. [pdf]
  • Walsh, J. 2012. Patterns of population structure and productivity in Saltmarsh Sparrows. MS Thesis. University of New Hampshire, Durham, NH. [pdf]
  • Walsh, J. 2015. Hybrid zone dynamics between saltmarsh (Ammodramus caudacutus) and Nelson’s (A. nelsoni) sparrows.  PhD Dissertation. University of New Hampshire, Durham, NH. [pdf]
  • Wiest, WA. 2010. Development of avian metrics to monitor salt marsh integrity. MS Thesis. University of Delaware, Newark, DE. [pdf]
  • Wiest, WA. 2015. Tidal marsh bird conservation in the Northeast, USA. Ph.D. Dissertation. University of Delaware, Newark, DE. [pdf]

Pre-SHARP publications on salt marshes from SHARP collaborators

  • Bayard, TS, and CS Elphick. 2010a. How area sensitivity in birds is studied. Conservation Biology 24:938-947. [pdf]
  • Bayard, TS, and CS Elphick. 2010b. Using spatial point-pattern assessment to understand the social and environmental mechanisms that drive avian habitat selection. Auk 127:485-494. [pdf]
  • Bayard, TS, and CS Elphick. 2012. Testing for conspecific attraction in an obligate saltmarsh bird: Can behavior be used to aid marsh restoration? Wetlands 32:521-529. [pdf]
  • Bayard, TS, and CS Elphick. 2011. Planning for sea-level rise: Quantifying patterns of Saltmarsh Sparrow (Ammodramus caudacutus) nest flooding under current sea-level conditions. Auk 128:393-403. [pdf]
  • Gjerdrum, C, CS Elphick, and M Rubega. 2005. Nest site selection and nesting success in saltmarsh breeding sparrows: the importance of nest habitat, timing, and study site differences. Condor 107:849-862. [pdf]
  • Gjerdrum, C, CS Elphick, and MA Rubega. 2008a. How well can we model numbers and productivity of saltmarsh sharp-tailed sparrows (Ammodramus caudacutus) using habitat features? Auk 125:608-617. [pdf]
  • Gjerdrum, C, K Sullivan-Wiley, E King, MA Rubega, and CS Elphick. 2008b. Egg and chick fates during tidal flooding of saltmarsh sharp-tailed sparrow nests. Condor 110:579-584. [pdf]
  • Greenberg, R, R Danner, B Olsen, and D Luther. 2012. High summer temperature explains bill size variation in salt marsh sparrows. Ecography 35:146-152. [pdf]
  • Greenberg, R, and B Olsen. 2010. Bill size and dimorphism in tidal-marsh sparrows: Island-like processes in a continental habitat. Ecology 91:2428-2436.  [pdf]
  • Greenberg, R, BJ Olsen, and MA Etterson. 2010. Patterns of seasonal abundance and social segregation in inland and Coastal Plain Swamp Sparrows in a delaware tidal marsh. Condor 112:159-167. [pdf]
  • Greenberg, R, C Elphick, JC Nordby, C Gjerdrum, H Spautz, G Shriver, B Schmeling, B Olsen, P Marra, N Nur, and M Winter. 2006. Flooding and predation: trade-offs in the nesting ecology of tidal-marsh sparrows.  In: Vertebrates of Tidal Marshes: Ecology, Evolution and Conservation (R Greenberg, S Droege, J Maldonado, and MV McDonald, eds). Studies in Avian Biology 32:96-109. [info here]
  • Hill, CE, C Gjerdrum, and CS Elphick. 2010. Extreme levels of multiple mating characterize the mating system of the Saltmarsh Sparrow (Ammodramus caudacutus). Auk 127:300-307. [pdf]
  • Hill, JM, and CS Elphick. 2011. Are grassland passerines especially susceptible to negative transmitter impacts? Wildlife Society Bulletin 35:362-367. [pdf]
  • Hill, JM, J Walsh, AI Kovach, and CS Elphick. 2013. Male-skewed sex ratio in Saltmarsh Sparrow nestlings. Condor 115:411-420. [pdf]
  • Humphries, S, CS Elphick, C Gjerdrum, and M  Rubega. 2007. Testing the function of the domed nests of Saltmarsh Sharp-tailed Sparrows. Journal of Field Ornithology 78:152-158. [link]
  • Kern, RA, WG Shriver, JL Bowman, LR Mitchell, and DL Bounds. 2012.  Seaside sparrow reproductive success in relation to prescribed fire. The Journal of Wildlife Management 76:932-939.  [pdf]
  • Lane, OP, KM O’Brien, DC Evers, TP Hodgman, A Major, N Pau, MJ Ducey, R Taylor, and D Perry. 2011. Mercury in breeding saltmarsh sparrows (Ammodramus caudacutus caudacutus). Ecotoxicology 20 (8): 1984–1991. [pdf]
  • Liu, IA, B Lohr, B Olsen, and R Greenberg. 2008. Macrogeographic vocal variation in subspecies of swamp sparrow. Condor 110:102-109.  [pdf]
  • Meiman, S, and CS Elphick. 2012. Evaluating habitat-association models for the Saltmarsh Sparrow. Condor 114:856-864. [pdf]
  • Meiman, S D Civco, K Holsinger, and CS Elphick. 2012. Comparing habitat models using ground-based and remote sensing data: saltmarsh sparrow presence versus nesting. Wetlands 32:725-736. [pdf]
  • Olsen, B, J Felch, R Greenberg, and J Walters. 2008a. Causes of reduced clutch size in a tidal marsh endemic. Oecologia 158:421-435. [pdf]
  • Olsen, BJ, R Greenberg, RC Fleischer, and JR Walters. 2008b. Extrapair paternity in the swamp sparrow, Melospiza georgiana: Male access or female preference? Behavioral Ecology & Sociobiology 63:285-294. [pdf]
  • Olsen, BJ, R Greenberg, IA Liu, JM Felch, and JR Walters. 2010. Interactions between sexual and natural selection on the evolution of a plumage badge. Evolutionary Ecology 24:731-748. [pdf]
  • Pepper, MA, and WG Shriver. 2010. Effects of open marsh water management on the reproductive success and nesting ecology of seaside sparrows in tidal marshes. Waterbirds 33:381-388. [pdf]
  • Shriver, WG, TP Hodgman, JP Gibbs, and PD Vickery. 2010. Home range sizes and habitat use of Nelson’s and Saltmarsh sparrows. Wilson Journal of Ornithology 122:340-345. [pdf]
  • Shriver, WG, PD Vickery, TP Hodgman, and JP Gibbs. 2007. Flood tides affect breeding ecology of two sympatric sharp-tailed sparrows. Auk 124:552-560. [pdf]
  • Walsh, J, AI Kovach, OP Lane, KM O’Brien, and KJ Babbitt. 2011. Genetic barcode rflp analysis of the Nelson’s and Saltmarsh sparrow hybrid zone. Wilson Journal of Ornithology 123:316-322. [pdf]
  • Walsh, J, AI Kovach, KJ Babbitt, and KM O’Brien. 2012. Fine-scale population structure and asymmetrical dispersal in an obligate salt-marsh passerine, the Saltmarsh Sparrow (Ammodramus caudacutus). Auk 129:247–258. [pdf]
  • Warner, SE, WG Shriver, MA Pepper, and RJ Taylor. 2010. Mercury concentrations in tidal marsh sparrows and their use as bioindicators in delaware bay, USA. Environmental Monitoring and Assessment 171:671-679. [pdf]

For complete lists of publications see the webpages of individual SHARP PI’s:
 Elphick,  KovachOlsenShriver, Cohen